View source for Biomes

<div class="map-thumb"><div>[[File:Biome Map.png|frameless|1066x1066px|class=no-pixel|link=Biomes map]]</div>[[Biomes map]]</div>

'''Biomes''' are the different areas within [[Dead Cells]] which players must progress through.<ref>[https://twinoid.com/en/article/5694/the-development-of-dead-cells-is-really-moving-on The development of Dead Cells is really moving on.] ''Twinoid.'' December 23, 2016.</ref> The base game currently has 15 main biomes and 4 boss biomes while the DLCs add another 9 main biomes and 8 boss biomes.<ref>[http://steamcommunity.com/games/588650/announcements/detail/1296319532119654152 Dead Cells is AVAILABLE NOW!] ''Steam.'' May 10, 2017.</ref> The order in which biomes are encountered, and the general location of their links, are predetermined but they are divided into levels and only one biome of each level can be accessed in a run which gives players a lot of flexibility and change each run.

Within each run, biomes are procedurally generated within the confines of that biome's set parameters. This puts the focus of gameplay on a combination of on the moment adaptation and memorization, with each biome having a distinct feel.<ref>[http://www.gamasutra.com/blogs/SebastienBENARD/20170329/294642/Building_the_Level_Design_of_a_procedurally_generated_Metroidvania_a_hybrid_approach.php Building the Level Design of a procedurally generated Metroidvania: a hybrid approach], ''Gamasutra''. March 29, 2017.</ref>

Each biome also has a set of [[enemies]] that the player will encounter, some of them are unique to a specific biome. Higher difficulties change the enemy distribution up.

== Scaling and difficulty ==
Each tier and biome has their own scaling of enemy and trap stats with tiers which operate somewhat differently compared to scaling of player damage and health with [[Stats]]. The exact formula used for scaling is not known at the moment but the tiers used for these are, meaning we can determine which biomes are easier and which are harder.

== Shops and treasures ==
Excluding boss biomes, each biome always contains combinations of shops and treasures. The majority of these are predetermined but their type might be random, shops might be either be a weapons or skills shop and treasure rooms could contain treasure chests, linked altars or very rarely an additional cursed chest. Their locations are always random except for the fact they will always have the same access requirements.

Playing on higher difficulty using collected [[Boss Stem Cells]] grants access to boss cell doors which can contain more shops, treasure rooms, or even new exits. 

== Access and exits ==
Each biome has exits leading to the biomes of the next level. Some of these can be accessed without any requirement but most are blocked off by rune requirement, which need to be found and unlocked first. The locations of exits in a biome is mostly random, with some exceptions, but are always at the end of hallways or level chunks. They are also seperated, meaning that if there are multiple exits a biome splits up creating different paths depending on the biome.

Additional higher difficulties also unlock new exits allowing new paths.

== [[Passage]] ==
After exiting each biome, the player will enter a [[Passage]]. Here, a player can turn in [[blueprints]], spend cells, enter [[time, killstreak and no-hit doors]] if the requirements are met, re-roll and upgrade [[gear]], choose and change [[mutations]] and replenish their health and health potions. Passages are different depending on which biome is exited and entered, with some of them having secrets.

After exiting a boss biome, the player will enter a special passage which also holds the [[Blacksmith]] and a no-hit door.

== Map ==
The player can open a biome map that will show everything the player has explored in the biome. Shown on the map are activated teleporters, shops and treasure rooms. The position of the player is marked with a flaming head, but this is not shown when inside a passage.

The player can also switch to a world map view, where discovered biomes are shown in color with their names, and undiscovered biomes are shown in gray without their name. A biome counts as discovered when it has been visited once, while an undiscovered biome is shown the moment a biome that can exit into that biome is entered.

Highlighting a discovered biome will show previously travelled paths between it and biomes of the previous and next tier, while the path used in the current run is indicated by a gray line. Furthermore, the world map will also show the next biome that has the [[Currency#Biome Incentive|incentivized biome bonus]].

== First stage ==
=== [[Prisoners' Quarters]] ===
[[File:Prisoners' Quarters.png|center|border|657x180px|class=res-img|frameless|link=The Prisoners' Quarters]]
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* '''Enemy Tier:''' 1 - 3
* '''Gear Level:''' 1
* '''Scrolls:''' 2 Power
* '''Enemies:''' [[Zombie]]s, [[Shieldbearer]]s, [[Grenadier]]s, [[Undead Archer]]s
**[[File:BSCI Icon.png|link=Boss Stem Cells]] [[Rancid Rat]]s, [[Knife Thrower]]s (replaces Undead Archers)
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Inquisitor]]s, [[Demolisher|Demolishers ,]]  [[Oven Knight]]s
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s
* '''Exits:''' [[Promenade of the Condemned]]
** {{Rune|vine}} [[Toxic Sewers]]
** {{Rune|tele}} [[Dilapidated Arboretum]]{{DLC|TBS}}
** [[Castle Outskirts]]{{DLC|RtC}}
* '''Doors:''' 
** '''Cavern Key Door''' - The big door at the start of the biome. Contains the {{Pickup|cavern key|1}}{{DLC|RotG}} and {{Rune|hom|1}} is required to get it. ([[The Hand of the King]] must have been beaten at least once)
* '''Keys:'''
** '''{{Pickup|cavern key|1}}:'''{{DLC|RotG}} Gives access to the [[Cavern]].{{DLC|RotG}}
** '''{{Pickup|dilapidated arboretum key|1}}:'''{{DLC|TBS}} Gives access to the [[Dilapidated Arboretum]].{{DLC|TBS}} It’s near the exit to it.
|}

== Second stages ==
=== [[Promenade of the Condemned]] ===
[[File:Promenade of the Condemned.png|center|border|657x180px|class=res-img|frameless|link=Promenade of the Condemned]]
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* '''Runes Needed to Access:''' None
* '''Enemy Tier:''' 3 - 7
* '''Gear Level:''' 2
* '''Scrolls:''' 1 Power, 1 extra power (1+ [[BSC]]), 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Zombie]]s, [[Grenadier]]s, [[Runner]]s, [[Bat]]s, [[Protector]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Kamikaze]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Dark Tracker]]s
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Bombardier]]s (Replaces Grenadiers)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s (Replaces Dark Trackers)
* '''Exits:''' [[Ramparts]] 
** {{Rune|spider}} [[Prison Depths]]
** {{Rune|tele}} [[Ossuary]]
** {{Rune|tele}} [[Morass of the Banished]]{{DLC|TBS}}
* '''Timed Door:''' 2 minutes ({{Item|Assault Shield}} blueprint)
* '''Hazards:''' Spikes, spiked flails
* '''Boss Cell Doors:''' 
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Shops|Weapon/Skill shop]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Objects#Cells_Vats|Cell Vat]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
* '''Keys:'''
** '''{{Pickup|Gardener's Key|1}}:''' There are 3 of them, give access to the blueprint of {{Item|Explosive Crossbow}}; also give access to 3 {{Pickup| Moonflower Key|1}}s when 3BSC or more are active.
* '''Runes:''' {{Rune|vin|1}}
|}

=== [[Toxic Sewers]] ===
[[File:Toxic Sewers.png|center|border|657x180px|class=res-img|frameless|link=Toxic Sewers]]
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* '''Runes Needed to Access:''' {{Rune|vin|1}}
* '''Enemy Tier:''' 4 - 7
* '''Gear Level:''' 3
* '''Scrolls:''' 1 Power, 1 extra power (1+ [[BSC]]), 2 Dual Scrolls, 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Zombie]]s, [[Scorpion]]s, [[Disgusting Worm]]s, [[Undead Archer]]s, [[Kamikaze]]s, [[Rancid Rat]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Grenadier]]s (Replaces Undead Archers)
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Buzzcutter]]s
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s (Replaces Zombies)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Festering Zombie]]s (Replaces Grenadiers), [[Corpse Worm]]s (spawned from Festering Zombies)
* '''Exits:''' [[Ramparts]]
** {{Rune|spi}} [[Corrupted Prison]]
** {{Rune|ram}} [[Ancient Sewers]]
* '''Timed Door:''' 2 minutes ({{Mutation|Frenzy}} blueprint)
* '''Hazards:''' Toxic pools
* '''Boss Cell Doors:''' 
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
* '''Runes:''' {{Rune|tel|1}}
|}

=== [[Dilapidated Arboretum]] ===
[[File:Dilapidated Arboretum.png|center|border|657x180px|class=res-img|frameless|link=Dilapidated Arboretum]]
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* '''Runes Needed to Access:''' {{Rune|tel|1}}
* '''Enemy Tier:''' 3 - 6
* '''Gear Level:''' 2
* '''Scrolls:''' 1 Power, 1 extra power (1+ [[BSC]]), 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Yeeter]]s,{{DLC|TBS}} [[Jerkshroom]]s,{{DLC|TBS}} [[Thorny|Thornies]], [[Zombie]]s, [[Bat]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Impaler (Enemy)|Impalers]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Kamikaze]]s (Replaces Bats)
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s (Replaces Zombies)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Buzzcutter]]s
* '''Exits:''' [[Morass of the Banished]],{{DLC|TBS}} [[Ramparts]]
** {{Rune|spider}} [[Prison Depths]]
* '''Timed Door:''' 2 minutes
* '''Hazards:''' Spikes, carnivorous plants
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Shops|Weapon/Skill shop]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Objects#Cells_Vats|Cell Vat]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
|}

=== [[Castle's Outskirts]] ===
[[File:Castle's Outskirts.png|center|border|657x180px|class=res-img|frameless|link=Castle's Outskirts]]
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* '''Entrances:''' [[Prisoners' Quarters]]
* '''Enemy Tier:''' 4 - 7
* '''Gear Level:''' II
* '''Scrolls:''' 1 Scroll of Power, 1 extra power (1+ [[BSC]]), 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Werewolf]]{{DLC|RTC}}, [[Merman]]{{DLC|RTC}}, [[Harpy]]{{DLC|RTC}}, [[Throw Master]]{{DLC|RTC}}, [[Vampire Bat]]{{DLC|RTC}}, [[Zombie]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Armor Knight]]{{DLC|RTC}}, [[Dark Tracker]], [[Demolisher]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Dire Werewolf]]{{DLC|RTC}} (Replaces Werewolf)
* '''Exits:''' [[Dracula's Castle]]{{DLC|RTC}}
* '''Hazards:''' 
* '''Boss Cell Doors:''' 
** [[File:BSCII Icon.png|link=Boss Stem Cells]] Skill or weapon Shop
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] Treasure Chest
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] Two choice chained items altar
* '''Keys:'''
** '''{{Pickup|Ribonned key|1}}:'''{{DLC|RTC}} Used to open Maria's Cell.
|}

== Optional stages ==
=== [[Prison Depths]] ===
[[File:Prison Depths.png|center|border|657x180px|class=res-img|frameless|link=Prison Depths]]
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* '''Entrances:''' [[Promenade of the Condemned]], [[Dilapidated Arboretum]]{{DLC|TBS}}
** {{Rune|spi|1}} required for all entrances.
* '''Enemy Tier:''' 8 - 9
* '''Gear Level:''' 3
* '''Scrolls:''' 1 Power (contained in a [[Cursed chest]])
* '''Enemies:''' [[Zombie]]s, [[Undead Archer]]s, [[Slasher]]s, [[Masker]]s, [[Hammer]]s, [[Sewer Fly|Sewer Flies]] (spawned from Hammers), [[Kamikaze]]s, [[Lacerator]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Knife Thrower]]s (Replaces Undead Archers)
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s (Replaces Zombies)
* '''Exits:''' [[Ossuary]], [[Morass of the Banished]]{{DLC|TBS}}
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Ancient Sewers]]
* '''Hazards:''' Spikes, spiked flails
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Ancient Sewers]] exit
* '''Keys:'''
** '''{{Pickup|Iron Cells Key|1}}:''' Give access to 1 of 2 treasure rooms in the end of the biome, each room contains two chained items.

Can only be accessed via the [[Promenade of the Condemned]] and [[Dilapidated Arboretum]].{{DLC|TBS}} This biome acts as an optional stage between the second and third stages.
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=== [[Corrupted Prison]] ===
[[File:Corrupted Prison.png|center|border|657x180px|class=res-img|frameless|link=Corrupted Confinement]]
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* '''Entrance:''' [[Toxic Sewers]] ({{Rune|spi|1}} required)
* '''Enemy Tier:''' 8 - 9
* '''Gear Level:''' 3
* '''Scrolls:''' 1 Power (contained in a [[Cursed chest]])
* '''Enemies:''' [[Slammer]]s, [[Shocker]]s, [[Bat]]s, [[Grenadier]]s, [[Zombie]]s, [[Rancid Rat]]s, [[Toxic Miasma]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Protector]]s (Replaces Shockers)
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Weirded Warrior]]s, [[Dark Tracker]]s (Replaces Zombies)
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Inquisitor]]s (Replaces Grenadiers)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Bomber]]s (Replaces Dark Trackers)
* '''Exits:''' [[Ancient Sewers]]
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Ramparts]]
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Ramparts]] exit
* '''Keys:'''
** '''{{Pickup|Iron Cells Key|1}}:''' Give access to 1 of 2 treasure rooms in the end of the biome, each room contains two chained items.

Can only be accessed via the [[Toxic Sewers]]. This biome acts as an optional stage between the second and third stages.
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== Third stages ==
=== [[Ramparts]] ===
[[File:Ramparts.png|center|border|657x180px|class=res-img|frameless|link=Ramparts]]
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* '''Entrances:''' [[Toxic Sewers]], [[Promenade of the Condemned]], [[Dilapidated Arboretum]],{{DLC|TBS}} [[Corrupted Prison]] (1+ [[BSC]])
* '''Enemy Tier:''' 7 - 13
* '''Gear Level:''' 3
* '''Scrolls:''' 3 Power, 1 extra Power (2+ [[BSC]]), 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Zombie]]s, [[Undead Archer]]s, [[Buzzcutter]]s, [[Inquisitor]]s, [[Sweeper]]s, [[Shieldbearer]]s
**[[File:BSCI Icon.png|link=Boss Stem Cells]] [[Weirded Warrior]]s, [[Slasher]]s, [[Demolisher]]s (replaces Undead Archers)
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Oven Knight]]s (Replaces Shieldbearers)
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager|Rampagers]] (Replaces Zombies)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Cannibal]]s (Replaces Slashers)
* '''Exits:''' [[Black Bridge]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Insufferable Crypt]]
* '''Timed Door:''' 8 minutes
* '''Hazards:''' Void
* '''Boss Cell Doors:''' 
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Shops|Food shop]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]], [[Insufferable Crypt]] exit
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
* '''Keys:'''
** '''{{Pickup|Moonflower Key|1}}:''' The first one. Hidden behind some foliage.
* '''Runes:''' {{Rune|custom|1}}
|}

=== [[Ancient Sewers]] ===
[[File:Ancient Sewers.png|center|border|657x180px|class=res-img|frameless|link=Ancient Sewers]]
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* '''Entrances:''' [[Toxic Sewers]] ({{Rune|ram|1}} required), [[Corrupted Prison]], [[Prison Depths]] (1+ [[BSC]])
* '''Enemy Tier:''' 6 - 13
* '''Gear Level:''' 4
* '''Scrolls:''' 3 Power, 1 extra Power (2+ [[BSC]]), 2 Dual Scrolls, 3 [[Scroll Fragments]] (3 [[BSC]]), 5 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Zombie]]s, [[Festering Zombie]]s, [[Corpse Worm]]s (spawned from Festering Zombies), [[Shieldbearer]]s, [[Disgusting Worm]]s, [[Kamikaze]]s, [[Impaler (Enemy)|Impalers]], [[Sewer's Tentacle]]s
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s (Replaces Zombies)
* '''Exit:''' [[Insufferable Crypt]]
* '''Hazards:''' Toxic pools, spikes, spiked flails
* '''Boss Cell Doors:''' 
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Objects#Chained_Items|Chained items]]
* '''Keys:'''
**'''{{Pickup|Old Service Key|1}}:''' 3 in total, used to access 3 of the 8 rewards at the end of the biome. Dropped by random enemies.
|}

=== [[Ossuary]] ===
[[File:Ossuary.png|center|border|657x180px|class=res-img|frameless|link=Ossuary]]
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* '''Entrances:''' [[Promenade of the Condemned]] ({{Rune|tel|1}} required), [[Prison Depths]]
* '''Enemy Tier:''' 7 - 13
* '''Gear Level:''' 3
* '''Scrolls:''' 2 Power (+1 in a guaranteed [[Cursed chest]]), 1 extra Power (2+ [[BSC]]), 2 Dual Scrolls, 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Zombie]]s, [[Grenadier]]s, [[Slasher]]s, [[Shocker]]s, [[Thorny|Thornies]], [[Spawner]]s, [[Corpse Juice]]s (spawned from Spawners)
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Inquisitor]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Bomber]]s (Replaces Grenadiers)
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Dark Tracker]]s (Replaces Zombies)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Bombardier]]s (Replaces Bombers)
* '''Exit:''' [[Black Bridge]]
* '''Timed Door:''' 8 minutes ({{Item|Marksman's Bow}} blueprint)
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Shops|Food shop]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
* '''Runes:''' {{Rune|ram|1}}
|}

=== [[Morass of the Banished]] === 
[[File:Morass of the Banished.png|center|border|657x180px|class=res-img|frameless|link=Morass of the Banished]]
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* '''Entrances:''' [[Dilapidated Arboretum]],{{DLC|TBS}} [[Promenade of the Condemned]] ({{Rune|tel|1}} required), [[Prison Depths]]
* '''Enemy Tier:''' 6 - 12
* '''Gear Level:''' 3
* '''Scrolls:''' 2 Power (+1 in a guaranteed [[Cursed chest]]), 1 extra Power (2+ [[BSC]]), 2 Dual Scrolls, 2 [[Scroll Fragments]] (3 [[BSC]]), 4 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Banished]],{{DLC|TBS}} [[Blowgunner]]s{{DLC|TBS}}, [[Giant Tick]]s{{DLC|TBS}}, [[Weaver Worm]]s, [[Cleaver (Enemy)|Cleavers]], [[Slasher]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Dark Tracker]]s
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s (Replaces Dark Trackers)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Masker]]s
* '''Exit:''' [[Nest]]{{DLC|TBS}}
* '''Boss Cell Doors:'''
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Shops|Food shop]]
|}

=== [[Dracula's Castle]] === 
[[File:Dracula's Castle.png|center|border|657x180px|class=res-img|frameless|link=Dracula's Castle]]
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* '''Entrances:''' [[Castle's Outskirts]]
* '''Enemy Tier:''' 6 - 13
* '''Gear Level:''' IV
* '''Scrolls:''' 3 Power (+1 in a guaranteed [[Cursed chest]]), 1 extra Power (2+ [[BSC]]), 2 Dual Scrolls, 3 [[Scroll Fragments]] (3 [[BSC]]), 4 [[Scroll Fragments]] (4/5 [[BSC]]) 
* '''Enemies:''' [[Harpy]]{{DLC|RTC}}, [[Axe Armor]]{{DLC|RTC}}, [[Buer]]{{DLC|RTC}}, [[Armor Knight]]{{DLC|RTC}}, [[Throw Master]]{{DLC|RTC}},  [[Werewolf]]{{DLC|RTC}} 
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Knife Thrower]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Dire Werewolf]]{{DLC|RTC}} (Replaces Werewolf), [[Guardian Knight]]
* '''Exit:''' [[Defiled Necropolis]]
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] Treasure Chest
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Shops|Food shop]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] Treasure Chest
|}

== First bosses ==
=== [[Black Bridge]] ===
[[File:Black Bridge.png|center|border|657x180px|class=res-img|frameless|link=Black Bridge]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Ossuary]], [[Ramparts]]
* '''Enemies:''' [[The Concierge]]
* '''Enemy Tier:''' 13
* '''Gear Level:''' 3
* '''Scrolls:''' 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Exits:''' [[Stilt Village]], [[Fractured Shrines]]{{DLC|FF}}
** {{Rune|spider}} [[Slumbering Sanctuary]]
* '''Timed Door (after the fight):''' 15 minutes ({{Item|Root Grenade}} blueprint)
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Runes:''' {{Rune|challenge|1}}
|}

=== [[Insufferable Crypt]] ===
[[File:Insufferable Crypt.png|center|border|657x180px|class=res-img|frameless|link=Insufferable Crypt]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Ancient Sewers]], [[Ramparts]] (3+ [[BSC]])
* '''Enemies:''' [[Conjunctivius]], Conjunctivius's Tentacles
* '''Enemy Tier:''' 11
* '''Gear Level:''' 3
* '''Scrolls:''' 3 [[Scroll Fragments]] (3 [[BSC]]), 5 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Timed Door (after the fight):''' 19:30 minutes (only if going to the [[Graveyard]])
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Exits:''' [[Slumbering Sanctuary]]
** {{Rune|spider}} [[Graveyard]]
* '''Hazards:''' Spikes
|}

=== [[Nest]] ===
[[File:Nest.png|center|border|657x180px|class=res-img|frameless|link=Nest]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrance:''' [[Morass of the Banished]]{{DLC|TBS}}
* '''Enemies:''' [[Mama Tick]]{{DLC|TBS}}
* '''Enemy Tier:''' 12
* '''Gear Level:''' 3
* '''Scrolls:''' 3 [[Scroll Fragments]] (3 [[BSC]]), 4 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Timed Door (after the fight):''' 15 minutes
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Exits:''' [[Stilt Village]], [[Fractured Shrines]]{{DLC|FF}}
** {{Rune|spider}} [[Graveyard]]
|}

=== [[Defiled Necropolis]] ===
[[File:Defiled Necropolis.png|center|border|657x180px|class=res-img|frameless|link=Defiled Necropolis]]
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* '''Entrance:''' Depth 3 [[Dracula's Castle]]{{DLC|RTC}}
* '''Enemies:''' [[Death]]{{DLC|RTC}}
* '''Enemy Tier:''' 13
* '''Gear Level:''' V
* '''Scrolls:''' 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Timed Door (after the fight):''' 15 minutes
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Exits:''' [[Slumbering Sanctuary]], [[Stilt Village]], [[Fractured Shrines]]{{DLC|FF}}
|}

== Fourth stages ==
=== [[Stilt Village]] ===
[[File:Stilt Village.png|center|border|657x180px|class=res-img|frameless|link=Stilt Village]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Black Bridge]], [[Nest]]{{DLC|TBS}}
* '''Enemy Tier:''' 14 - 18
* '''Gear Level:''' 4
* '''Scrolls:''' 3 Power, 1 extra power (3+ [[BSC]]), 1 Dual Scroll, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Festering Zombie]]s, [[Corpse Worm]]s (spawned from Festering Zombies), [[Weaver Worm]]s, [[Pirate Captain]]s, [[Zombie]]s, [[Kamikaze]]s, [[Weirded Warrior]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Demolisher]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Bomber]]s
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s (Replaces Zombies)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s
* '''Exits:''' [[Clock Tower]], [[Undying Shores]]{{DLC|FF}} (requires reaching the Undying Shores from the [[Fractured Shrines]]{{DLC|FF}} with the {{Outfit|Cultist Outfit}}{{DLC|FF}} at least once)
** {{Rune|teleport}} [[Forgotten Sepulcher]]
* '''Boss Cell Doors:''' 
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
* '''Keys:'''
**'''{{Pickup|Village Key|1}}:''' 2 keys can be found in 2 key rooms in the biome, which lead to exits to [[Forgotten Sepulcher]], [[Undying Shores]]{{DLC|FF}} or the [[Clock Tower]], or blueprint of {{Item|Parry Shield}}.
|}

=== [[Slumbering Sanctuary]] ===
[[File:Slumbering Sanctuary.png|center|border|657x180px|class=res-img|frameless|link=Slumbering Sanctuary]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Black Bridge]] ({{Rune|spi|1}} required), [[Insufferable Crypt]]
* '''Enemy Tier:''' 15 - 20
* '''Gear Level:''' 4
* '''Scrolls:''' 2 Power (+1 in a guaranteed [[Cursed chest]]), 1 extra power (3+ [[BSC]]), 1 Dual Scroll, 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' 
** '''Before activating AT switch:''' [[Zombie]]s, [[Catcher]]s, [[Masker]]s, [[Caster]]s
*** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Cannibal]]s
** '''After activating AT switch:''' [[Grenadier]]s, [[Inquisitor]]s, [[Protector]]s, [[Kamikaze]]s, [[Golem]]s, [[Oven Knight]]s, [[Dancer]]s
*** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Bombardier]]s (Replaces Grenadiers)
*** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s
*** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Demon]]s (Replaces Inquisitors)
* '''Exits:''' [[Clock Tower]], [[Forgotten Sepulcher]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Cavern]]{{DLC|RotG}} ([[The Giant]]{{DLC|RotG}} has to have been killed at least once)
* '''Timed Door:''' 26 minutes
* '''Hazards:''' Spikes, spiked flails, sawblades
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Objects#chained items|Chained items]], [[Cavern]]{{DLC|RotG}} exit
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
* '''Runes:''' {{Rune|spider|1}}
|}

=== [[Graveyard]] ===
[[File:Graveyard.png|center|border|657x180px|class=res-img|frameless|link=Graveyard]]
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* '''Entrances:''' [[Insufferable Crypt]], [[Nest]]{{DLC|TBS}}
** {{Rune|spi|1}} required for all entrances.
* '''Enemy Tier:''' 14 - 19
* '''Gear Level:''' 4
* '''Scrolls:''' 2 Power (+1 in a guaranteed [[Cursed chest]]), 1 extra power (3+ [[BSC]]), 1 Dual Scroll, 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Swarm Zombie]]s, [[Corpse Fly|Corpse Flies]] (spawned from Swarm Zombies), [[Inquisitor]]s, [[Catcher]]s, [[Bat]]s, [[Kamikaze]]s, [[Masker]]s, [[Rancid Rat]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Protector]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Bomber]]s (Replaces Bats)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Cannibal]]s
* '''Exits:''' [[Forgotten Sepulcher]], [[Cavern]]{{DLC|RotG}} (requires the {{Pickup|Cavern Key}}{{DLC|RotG}} on first exit), [[Undying Shores]]{{DLC|FF}}
* '''Timed Door:''' 19 minutes, 30 seconds
* '''Hazards:''' Spikes, spiked flails
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Objects#Chained Items|Chained items]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Shops|Food shop]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Objects#Cells Vats|Cells vat]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Objects#Chained Items|Chained items]]
* '''Keys:'''
** '''{{Pickup|Graveyard Key|1}}:''' Give access to the bottom half of the biome.
** '''{{Pickup|Architect's Key|1}}:''' Give access to blueprints of {{Mutation|Parting Gift}} and [[The Collector#General improvements|Merchandise Categories]]. Hidden in a random Wall Secret.
** '''{{Pickup|Moonflower Key|1}}:''' The second one. Hidden behind some foliage.
|}

=== [[Fractured Shrines]] ===
[[File:Fractured Shrines.png|center|border|657x180px|class=res-img|frameless|link=Fractured Shrines]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Black Bridge]], [[Nest]]{{DLC|TBS}}
* '''Enemy Tier:''' 12 - 16
* '''Gear Level:''' 4
* '''Scrolls:''' 1 Power (+1 in a guaranteed [[Cursed chest]]), 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Myopic Crow]]s,{{DLC|FF}} [[Stone Warden]]s,{{DLC|FF}} [[Cold Blooded Guardian]]s,{{DLC|FF}} [[Slasher]]s, [[inquisitor]]s, [[Zombie]]s, [[Grenadier]]s, [[Undead Archer]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Shocker]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Bombardier]]s (Replaces Grenadiers)
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Bomber]]s (Replaces Zombies)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Catcher]]s (Replaces Undead Archers)
* '''Exits:''' [[Undying Shores]]{{DLC|FF}} (requires the {{Outfit|Cultist Outfit}}{{DLC|FF}} on first exit), [[Clock Tower]]
** {{Rune|teleport}} [[Forgotten Sepulcher]] (in the passage to the Clock Tower)
* '''Hazards:''' Pits, rotating axe, swinging log, [[Myopic Crow]]s{{DLC|FF}}
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
|}

== Fifth stages ==
=== [[Clock Tower]] ===
[[File:Clock Tower.png|center|border|657x180px|class=res-img|frameless|link=Clock Tower]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Stilt Village]], [[Slumbering Sanctuary]], [[Fractured Shrines]]{{DLC|TQatS}}
* '''Enemy Tier:''' 19 - 23
* '''Gear Level:''' 5
* '''Scrolls:''' 4 Power, 1 extra power (4+ [[BSC]]), 2 Dual Scrolls, 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Dark Tracker]]s, [[Cannibal]]s, [[Bombardier]]s, [[Undead Archer]]s, [[Automaton]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Oven Knight]]s
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Demolisher]]s (replace [[Undead Archer]]s)
* '''Exit:''' [[Clock Room]]
* '''Hazards:''' Spikes, spiked flails
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Shops|Weapon/Skill shop]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Objects#Cells Vat|Cell vat]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]] 
* '''Keys:'''
** '''{{Pickup|Clockmaker's Key|1}}:''' Gives access to the [[Clock Room]]. Located in one of the three or four towers.
** '''{{Pickup|Bell Tower Key|1}}:''' Gives access to blueprint of {{Item|Punishment}}. Look for the 4 Bells and hit them once from low to high pitch.
|}

=== [[Forgotten Sepulcher]] ===
[[File:Forgotten Sepulcher.png|center|border|657x180px|class=res-img|frameless|link=Forgotten Sepulcher]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Stilt Village]] ({{Rune|tel|1}} required), [[Slumbering Sanctuary]], [[Graveyard]], [[Fractured Shrines]]{{DLC|FF}} ({{Rune|tel|1}} required)
* '''Enemy Tier:''' 19 - 23
* '''Gear Level:''' 5
* '''Scrolls:''' 3 Power (+1 in a guaranteed [[Cursed chest]]), 1 extra power (4+ [[BSC]]), 2 Dual Scrolls, 3 [[Scroll Fragments]] (3 [[BSC]]), 4 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Dark Tracker]]s, [[Inquisitor]]s, [[Lacerator|Lacerators]], [[Cleaver (Enemy)|Cleavers]], [[Kamikaze]]s, [[Shocker]]s, [[Corpulent Zombie]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Knife Thrower]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Shieldbearer]]s, [[Weirded Warrior]]s
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s (Replaces Inquisitors)
* '''Exits:''' [[Clock Room]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Guardian's Haven]]{{DLC|RotG}} ([[The Giant]]{{DLC|RotG}} has to have been killed at least once)
* '''Timed Door:''' 26 minutes
* '''Hazards:''' Spikes, darkness
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Objects#Chained Items|Chained items]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Objects#Chained Items|Chained items]]
* '''Keys:'''
** '''{{Pickup|Crypt Key|1}}:''' There are 2 of them, each leading player further in the level.
** '''{{Pickup|Moonflower Key|1}}:''' The third one. Hidden behind some foliage.
* '''Runes:''' {{Rune|exp|1}}
|}

=== [[Cavern]] ===
[[File:Cavern.png|center|border|657x180px|class=res-img|frameless|link=Cavern]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Graveyard]] ({{Pickup|Cavern Key}}{{DLC|RotG}} required on first entry), [[Slumbering Sanctuary]] (2+ [[BSC]] and kill the [[Giant]]{{DLC|RotG}} at least once)
* '''Enemy Tier:''' 20 - 24
* '''Gear Level:''' 7
* '''Scrolls:''' 4 Power, 1 extra power (4+ [[BSC]]), 2 Dual Scrolls, 4 [[Scroll Fragments]] (3 [[BSC]]), 5 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Ground Shaker]]s,{{DLC|RotG}} [[Inquisitor]]s, [[Arbiter]]s,{{DLC|RotG}} [[Demon]]s, [[Lacerator]]s, [[Slammer]]s, [[Skeleton]]s{{DLC|RotG}}
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Oven Knight]]s
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s (Replace Lacerators)
* '''Exits:''' [[Guardian's Haven]],{{DLC|RotG}} [[Mausoleum]]{{DLC|FF}}
* '''Hazards:''' Pools of lava, lanterns, spikes, spiked flails, electric waves
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Objects#Chained Items|Chained items]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] {{Outfit|Festive Outfit}}
* '''Keys'''
** '''{{Pickup|Garland Key|1}}:'''{{DLC|RotG}} Gives access to the 4 [[BSC]] door which contains the blueprint of {{Outfit|Festive Outfit}}.{{DLC|RotG}} Look for a vine passage in the ceiling, often near a shop.
|}

=== [[Undying Shores]] ===
[[File:Undying Shores.png|center|border|657x180px|class=res-img|frameless|link=Undying Shores]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Fractured Shrines]]{{DLC|FF}} ({{Outfit|Cultist Outfit}}{{DLC|FF}} needs to be equipped on first entry), [[Stilt Village]], [[Graveyard]].
** The latter two entrances are available after the entrance in the [[Fractured Shrines]]{{DLC|FF}} has been unlocked.
* '''Enemy Tier:''' 19 - 23	
* '''Gear Level:''' 6
* '''Scrolls:''' 4 Power, 2 Dual Scrolls, 3 [[Scroll Fragments]] (3 [[BSC]]), 4 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Enemies:''' [[Apostate]]s,{{DLC|FF}} [[Failed Homunculus|Failed Homunculi]],{{DLC|FF}} [[Compulsive Gravedigger]]s,{{DLC|FF}} [[Clumsy Swordsman|Clumsy Swordsmen]],{{DLC|FF}} [[Dastardly Archer]]s,{{DLC|FF}} [[Grenadier]]s, [[Bat]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Knife Thrower]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Slammer]]s (Replace Bats)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Rampager]]s
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s
* '''Exit:''' [[Mausoleum]]{{DLC|FF}}
* '''Hazards:''' Pits
|}

== Second bosses ==
=== [[Clock Room]] ===
[[File:Clock Room.png|center|border|657x180px|class=res-img|frameless|link=Clock Room]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Clock Tower]], [[Forgotten Sepulcher]]
* '''Enemies:''' [[The Time Keeper]]
* '''Enemy Tier:''' 22
* '''Gear Level:''' 5
* '''Scrolls:''' 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Exits:''' [[High Peak Castle]], [[Derelict Distillery]] (requires meeting the [[Hand of the King]] at least once), [[Infested Shipwreck]]{{DLC|TQatS}}
|}

=== [[Guardian's Haven]] ===
[[File:Guardian's Haven.png|center|border|657x180px|class=res-img|frameless|link=Guardian's Haven]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Cavern]],{{DLC|RotG}} [[Forgotten Sepulcher]] (2+ [[BSC]] and kill the [[Giant]]{{DLC|RotG}} at least once)
* '''Enemies:''' [[The Giant]]{{DLC|RotG}}
* '''Enemy Tier:''' 21
* '''Gear Level:''' 7
* '''Scrolls:''' 3 [[Scroll Fragments]] (3 [[BSC]]), 4 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Exits:''' [[High Peak Castle]], [[Derelict Distillery]], [[Infested Shipwreck]],{{DLC|TQatS}} [[Throne Room]] (choosing this exit grants 2 Scrolls of Power)
* '''Hazards:''' Pools of lava
|}

=== [[Mausoleum]] ===
[[File:Mausoleum.png|center|border|657x180px|class=res-img|frameless|link=Mausoleum]]
{| class="wikitable mw-collapsible mw-collapsed" data-collapsetext="▲" data-expandtext="▼" style="text-align:left; width:100%; padding:1px; margin-left: auto; margin-right: auto;
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* '''Entrances:''' [[Undying Shores]],{{DLC|FF}} [[Cavern]]{{DLC|RotG}}
* '''Enemies:''' [[The Scarecrow]]{{DLC|FF}}
* '''Enemy Tier:''' 22
* '''Gear Level:''' 6
* '''Scrolls:''' 2 [[Scroll Fragments]] (3 [[BSC]]), 3 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Exits:''' [[High Peak Castle]], [[Derelict Distillery]], [[Infested Shipwreck]]{{DLC|TQatS}}
* '''Hazards:''' Explosive mushrooms
|}

== Sixth stages ==
=== [[High Peak Castle]] ===
[[File:High Peak Castle.png|center|border|657x180px|class=res-img|frameless|link=High Peak Castle]]
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* '''Entrances:''' [[Clock Room]], [[Mausoleum]],{{DLC|FF}} [[Guardian's Haven]]{{DLC|RotG}}
* '''Enemies:''' [[Guardian Knight]]s, [[Lancer]]s, [[Undead Archer]]s, [[Bombardier]]s, [[Royal Guard]]s
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Knife Thrower]]s (Replaces Undead Archers)
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Rampager]]s
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Demon]]s (Replaces Knife Throwers)
* '''Enemy Tier:''' 24 - 27
* '''Gear Level:''' 6
* '''Scrolls:''' 2 Power, 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Exit:''' [[Throne Room]]
* '''Hazards:''' Spikes, spiked flails
* '''Doors:''' 
** '''Castle Key Doors:''' They are 6 doors of 3 different colors, 2 doors each. Each 2 doors lead to an extra section of the biome, which offers an Elite enemy fight and a Castle Key as reward.
* '''Keys:'''
** '''{{Pickup|Castle Key|1}}:''' 3 in total. 2 are required to access [[Throne Room]], while the 3rd gives access to blueprint of {{Item|Boomerang}} and a Scroll of Power. Dropped by the three Elite enemies in each of the Castle Key doors.
|}

=== [[Derelict Distillery]] ===
[[File:Derelict Distillery.png|center|border|657x180px|class=res-img|frameless|link=Derelict Distillery]]
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* '''Entrances:''' [[Clock Room]], [[Mausoleum]],{{DLC|FF}} [[Guardian's Haven]]{{DLC|RotG}}
* '''Enemies:''' [[Living Barrel]]s, [[Infected Worker]]s, [[Demolisher]]s, [[Shieldbearer]]s, [[Rancid Rat]]s, [[Lacerator]]s, [[Spawner]]s, [[Corpse Juice]]s (spawned from Spawners)
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Bombardier]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Hammer]]s, [[Sewer Fly|Sewer Flies]] (spawned from Hammers), [[Oven Knight]]s (Replace Shieldbearers)
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Failed Experiment]]s
* '''Enemy Tier:''' 17 - 21
* '''Gear Level:''' 6
* '''Scrolls:''' 2 Power, 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Exits:''' [[Throne Room]], [[Lighthouse]]{{DLC|TQatS}}
* '''Hazards:''' Exploding barrels, Spikes
* '''Doors:''' There is a locked door blocking the exit to the [[Throne Room]] requiring the Distillery Key.
* '''Keys:'''
** '''{{Pickup|Distillery Key|1}}:''' Gives access to the Throne Room. Can be found behind a destructible wall, which can only be destroyed by deflecting a nearby exploding barrel or by using the {{Item|Barrel Launcher}}.
|}

=== [[Infested Shipwreck]] ===
[[File:Infested Shipwreck.png|center|border|657x180px|class=res-img|frameless|link=Infested Shipwreck]]
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* '''Entrances:''' [[Clock Room]], [[Mausoleum]],{{DLC|FF}} [[Guardian's Haven]]{{DLC|RotG}}
* '''Enemies:''' [[Mutineer]]s,{{DLC|TQatS}} [[Armored Shrimp]]s,{{DLC|TQatS}} [[Bombardier]]s, [[Rancid Rat]]s, [[Kamikaze]]s
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Pirate Captains]]s
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Rampager]]s
* '''Enemy Tier:''' 24 - 27
* '''Gear Level:''' 6
* '''Scrolls:''' 2 Power, 2 Dual Scrolls, 1 [[Scroll Fragment]] (3 [[BSC]]), 2 [[Scroll Fragments]] (4/5 [[BSC]])
* '''Exit:''' [[Lighthouse]]{{DLC|TQatS}}
* '''Hazards:''' Breakable platforms
* '''Boss Cell Doors:'''
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIII Icon.png|link=Boss Stem Cells]] [[Treasure chest]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] Chained items altar
* '''Keys:'''
** '''{{Pickup|Forked Key|1}}:''' Forked is used to unlock the loreroom where the [[Abyssal Trident]] can be found.
|}

=== [[Dracula's Castle]] === 
[[File:Dracula's Castle.png|center|border|657x180px|class=res-img|frameless|link=Dracula's Castle]]
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* '''Entrances:''' [[Clock Room]]
* '''Enemy Tier:''' 24  27
* '''Gear Level:''' VI
* '''Scrolls:''' 4 Power (+1 in a guaranteed [[Cursed chest]]), 1 extra Power (2+ [[BSC]]), 2 Dual Scrolls, 3 [[Scroll Fragments]] (3 [[BSC]]), 4 [[Scroll Fragments]] (4/5 [[BSC]]) 
* '''Enemies:''' [[Harpy]]{{DLC|RTC}}, [[Axe Armor]]{{DLC|RTC}}, [[Buer]]{{DLC|RTC}}, [[Armor Knight]]{{DLC|RTC}}, [[Throw Master]]{{DLC|RTC}},  [[Werewolf]]{{DLC|RTC}} 
** [[File:BSCI Icon.png|link=Boss Stem Cells]] [[Dire Werewolf]]{{DLC|RTC}} (Replaces Werewolf), [[Knife Thrower]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] [[Demon]]
* '''Exit:''' [[Master's Keep]]
* '''Boss Cell Doors:'''
** [[File:BSCI Icon.png|link=Boss Stem Cells]] Treasure Chest
** [[File:BSCII Icon.png|link=Boss Stem Cells]] [[Shops|Food shop]]
** [[File:BSCIV Icon.png|link=Boss Stem Cells]] Treasure Chest
|}

== Third bosses ==
=== [[Throne Room]] ===
[[File:Throne Room.png|center|border|657x180px|class=res-img|frameless|link=Throne Room]]
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* '''Entrances:''' [[High Peak Castle]], [[Derelict Distillery]], [[Guardian's Haven]]{{DLC|RotG}}
* '''Enemies:''' [[The Hand of the King]]
** ''Mobs summoned by the Hand of the King (2/5 will be Elites):'' [[Zombie]], [[Inquisitor]], [[Grenadier]], [[Lacerator]], [[Scorpion]], [[Disgusting Worm]], [[Cleaver (Enemy)|Cleaver]], [[Slasher]]
* '''Enemy Tier:''' 27
* '''Perfect Door (after the fight):''' Flawless fight ([[Legendary]] item)
* '''Exit:''' [[File:BSCV Icon.png|link=Boss Stem Cells]] [[Astrolab|Seventh stage]]{{DLC|RotG}}
* '''Hazards:''' Spikes
* '''Runes:''' {{Rune|hom|1}}
|}

=== [[Lighthouse]] ===
[[File:Lighthouse.png|center|border|657x180px|class=res-img|frameless|link=Lighthouse]]
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* '''Entrances:''' [[Infested Shipwreck]],{{DLC|TQatS}} [[Derelict Distillery]]
* '''Enemy Tier:''' 27
* '''Enemies:''' [[The Servants]]{{DLC|TQatS}}
* '''Perfect Door (after the fight):''' No
* '''Exit:''' [[The Crown]]{{DLC|TQatS}}
* '''Hazards:''' Bottomless pits, fire
|}

=== [[Master's Keep]] ===
[[File:Master's Keep.png|center|border|657x180px|class=res-img|frameless|link=Master's Keep]]
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* '''Entrances:''' [[Clock Tower]]
* '''Enemy Tier:''' 27
* '''Enemies:''' [[Dracula]] and [[Dracula - Final Form]]
* '''Perfect Door (after the fight):''' No
* '''Hazards:''' Bottomless pits
|}

== Seventh stage ==
{{SpoilerTag
|{{fake heading|sub=3|[[Astrolab]]}}
[[File:Astrolab.png|center|border|657x180px|class=res-img|frameless|link=Astrolab]]

* '''Entrance:''' [[Throne Room]] (5 [[BSC]])
* '''Enemy Tier:''' 37 - 46
* '''Gear Level:''' 10
* '''Scrolls:''' 2 Dual Scrolls
* '''Enemies:''' [[Bomber]]s, [[Defender]]s,{{DLC|RotG}} [[Failed Experiment]]s, [[Magistrate of Death|Magistrates of Death]],{{DLC|RotG}} [[Screaming Skull]]s{{DLC|RotG}} (spawned from Magistrates of Death), [[Slammer]]s, [[Librarian]]s{{DLC|RotG}}
* '''Exit:''' [[Observatory]]{{DLC|RotG}}
* '''Hazards:''' Electric waves, projectiles, spikes, void, pools of lava
* '''Keys:'''
** '''{{Pickup|Allen Key}}:'''{{DLC|RotG}} Opens the door to the final section of the biome. Dropped by an Elite [[Slammer]].
** '''{{Pickup|Elevator Key}}:'''{{DLC|RotG}} Opens the door before the elevator at the end of the biome. Dropped by an Elite [[Failed Experiment]].
** '''{{Pickup|Guardian's Key}}:'''{{DLC|RotG}} There are 2. Open the doors before the exit to [[Observatory]].{{DLC|RotG}} Dropped by 2 Elite [[Slammer]]s.
** '''{{Pickup|Apex Key}}:'''{{DLC|RotG}} Gives access time blueprint of {{Item|Sonic Carbine}}.{{DLC|RotG}} Found at the end of an obstacle course.
}}

== Fourth bosses ==
=== [[The Crown]] ===
[[File:The Crown.png|center|border|657x180px|class=res-img|frameless|link=The Crown]]
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* '''Entrance:''' [[Lighthouse]]{{DLC|TQatS}}
* '''Enemies:''' [[The Queen]]{{DLC|TQatS}}
* '''Enemy Tier:''' 27
|}

{{SpoilerTag
|{{fake heading|sub=3|[[Observatory]]}}
[[File:Observatory.png|center|border|657x180px|class=res-img|frameless|link=Observatory]]

* '''Entrance:''' [[Astrolab]]{{DLC|RotG}}
* '''Enemies:''' [[The Collector (5 BSC)|The Collector]]{{DLC|RotG}}
** ''Mobs summoned by the Collector:'' [[Grenadier]], [[Lancer]], [[Runner]], [[Undead Archer]], [[Cleaver (Enemy)|Cleaver]]
* '''Enemy Tier:''' 38
}}

== Special stages ==
=== [[The Bank]] ===
[[File:The Bank.png|center|border|657x180px|class=res-img|frameless|link=The Bank]]
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* '''Entrances:''' Any transition that does not lead to a boss.
* '''Enemies:''' [[Agitated Pickpocket]]s, [[Gold Gorger]]s, [[Golden Kamikaze]]s, [[Mimic]]s
** Enemies that spawn in this biome depends on what biomes it replaces.
* '''Enemy Tier:''' 
* '''Gear Level:''' 
* '''Scrolls:''' The Bank will contain one less scroll than the "best" biome that's being replaced, i.e. the one with the most scrolls. The lost scroll is replaced by a cursed chest.
* '''Exit:''' 
* '''Hazards:''' Spikes, spiked flails
* '''Doors:''' 
** '''Bank floors:''' There three floors in the bank that are blocked by locked doors. Each needs a pass of matching color to unlock.
* '''Keys:'''
** '''{{Pickup|Red pass|1}}:''' Unlocks the red door.
** '''{{Pickup|Blue pass|1}}:''' Unlocks the blue door.
** '''{{Pickup|Green pass|1}}:''' Unlocks the green door.
|}


== Unused stages ==
=== [[Repository of the Architects]] ===
An inaccessible stage that was a prototype for the [[Forgotten Sepulcher]], the entrance was behind a door in the Graveyard that required 5 boss cells to open. This door was impossible to open normally as it was removed before the Rise of the Giant DLC added the fifth boss stem cell. If accessed via modifying the boss cell counter with third-party tools, the stage could be seen with unique lighting very similar to what the Forgotten Sepulcher currently has, although it did not decay akin to how the darkness mechanic currently works. The only enemies present in the stage appeared to be [[Cannibal]]s. The enemy tier also appeared to be absurdly high, making them very hard to deal with due to their large HP and damage values. The majority of the stage was locked behind a door that required a key (simply named "Key") to open, which had no corresponding item ID and thus couldn't be spawned, even by third-party tools.

This stage is irrelevant to the {{Pickup|Architect's Key|1}}.

=== [[Pier]] ===
An old ending point of a run that persisted until the 1.0 update came around. It would be accessed after defeating the final boss of the game, which at first was [[Conjunctivius]], then the [[Time Keeper]], and finally, the [[Hand of the King]]. It used to have the [[NPCs#Unused NPCs|Fisherman NPC]] right next to a boat, who would impale the Beheaded with a tentacle to let the player start a new run. Later on, that was changed to a "Work in Progress" sign and a tube, through which the Beheaded would crawl to go back to the [[Prisoners' Quarters]]. Its assets have now been repurposed for the area leading to the [[Infested Shipwreck]].{{DLC|TQatS}}

== Gallery ==
<gallery widths="320" heights="180">
File:World Map.png|The world map partially explored.
File:Incentivized Biome.png|Incentivized biome pop-up. 
</gallery>

== References ==
<references />

{{Nav}}

[[Category:Gameplay]]
[[Category:Biomes]]